Symptoms:

You recently moved your WP (3.3.1) to a W2K R2 server, and now you can’t upload images. You are getting this error message:

Failed to upload due to an error:

The uploaded file could not be moved to ...

Solution:

Forget for a second about permissions on folder wp-contentuploads.

Instead, concentrate and fix permissions on folder wp_content.

For an inspirational discussion on this topic see:

http://wordpress.org/support/topic/file-upload-error-after-upgrading-to-33

All other tricks didn’t work for me. I would suggest this material as a base for a new article.